Martin Krarup Nielsen

2025

Australian guidelines for equine internal parasite management

Beasley, A., Abbas, G., Hughes, K., El-Hage, C., Jacobson, C., Bauquier, J., Wilkes, E., Carrigan, P., Cudmore, L., Hurley, J., Beveridge, I., Nielsen, M. & Jabbar, A., Apr 2025, In: Australian Veterinary Journal. 103, 4, p. 151-158

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Open Access

Ivermectin performance against equine strongylids: Efficacy, egg reappearance periods, and fecal egg counting method comparison

Nielsen, M. K., Bartholdy, I. D., Kristensen, K. S., Borrye, J. C., Meilvang, K. S. S., Rendtorff, C. E. K., Hjortflod, M. D., Fuglbjerg, V., Karlsson, M., Petersen, H. H., Toft, K., Thamsborg, S. M. & Pihl, T. H., Jun 2025, In: Veterinary Parasitology. 336, 110465.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

Profiling host- and parasite-derived miRNAs associated with Strongylus vulgaris infection in horses

Toft, K., Honoré, M. L., Ripley, N., Nielsen, M. K., Mardahl, M., Fromm, B., Hedberg-Alm, Y., Tydén, E., Nielsen, L. N., Nejsum, P., Thamsborg, S. M., Cirera, S. & Pihl, T. H., Feb 2025, In: Veterinary Parasitology. 334, 110379.

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Open Access

The microRNAome of Strongylus vulgaris larvae and their excretory/secretory products with identification of parasite-derived microRNAs in horse arterial tissue

Toft, K., Honoré, M. L., Ripley, N. E., Nielsen, M. K., Fromm, B., Mardahl, M., Nielsen, L. N., Nejsum, P., Thamsborg, S. M., Cirera, S. & Pihl, T. H., Jan 2025, In: International Journal for Parasitology. 55, 1, p. 45-58 14 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

1 Citation (Scopus)

2024

A diverse microbial community and common core microbiota associated with the gonad of female Parascaris spp.

Cain, J. L., Norris, J. K., Swan, M. P. & Nielsen, M. K., Jan 2024, In: Parasitology Research. 123, 1, 56.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

All that glitters is not gold: An interpretive framework for diagnostic test evaluation using Ascaris lumbricoides as a conceptual example

Denwood, M., Nielsen, S. S., Olsen, A., Jones, H. E., Coffeng, L. E., Landfried, G., Nielsen, M. K., Levecke, B., Thamsborg, S. M., Eusebi, P., Meletis, E., Kostoulas, P., Hartnack, S., Erkosar, B. & Toft, N., Sept 2024, In: PLOS Neglected Tropical Diseases . 18, 9, e0012481.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

A national survey of anthelmintic resistance in ascarid and strongylid nematodes in Australian Thoroughbred horses

Abbas, G., Ghafar, A., McConnell, E., Beasley, A., Bauquier, J., Wilkes, E. J. A., El-Hage, C., Carrigan, P., Cudmore, L., Hurley, J., Gauci, C. G., Beveridge, I., Ling, E., Jacobson, C., Stevenson, M. A., Nielsen, M. K., Hughes, K. J. & Jabbar, A., Apr 2024, In: International Journal for Parasitology: Drugs and Drug Resistance. 24, 100517.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

11 Citations (Scopus)

Bacillus thuringiensis Cry14A family proteins as novel anthelmintics against gastrointestinal nematode parasites

Hoang, D., Flanagan, K., Ding, Q., Cazeault, N. R., Li, H., Díaz-Valerio, S., Rus, F., Darfour, E. A., Kass, E., Petersson, K. H., Nielsen, M. K., Liesegang, H., Ostroff, G. R. & Aroian, R. V., Oct 2024, In: PLOS Neglected Tropical Diseases . 18, 10, e0012611.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

2 Citations (Scopus)

Evidence of tapeworm treatment failure on a Central Kentucky Thoroughbred farm

Finnerty, C. A., Bonometti, S., Ripley, N. E., Smith, M. A. & Nielsen, M. K., Nov 2024, In: Equine Veterinary Education. 36, 11, p. 579-585 7 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

4 Citations (Scopus)

First report of ivermectin resistance in cyathostomins (small strongyles) of horses in Argentina

Cooper, L. G., Benard, B. P., Nielsen, M. K., Caffe, G., Arroyo, F. & Anziani, O. S., Jul 2024, In: Veterinary Parasitology: Regional Studies and Reports. 52, 101046.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

1 Citation (Scopus)

Ivermectin performance in horses diagnosed with equine endocrine disorders

Nielsen, M. K., Finnerty, C. A., Ripley, N. E., Page, A. E., McClendon, M. E. & Adams, A. A., Jun 2024, In: Veterinary Parasitology. 328, 110182.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Performance of three techniques for diagnosing equine tapeworm infection

Anderson, H. C., Warner, S. F., Ripley, N. E. & Nielsen, M. K., Apr 2024, In: Veterinary Parasitology. 327, 110152.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

1 Citation (Scopus)

Protein profile of extracellular vesicles derived from adult Parascaris spp.

Manikantan, V., Ripley, N. E., Nielsen, M. K. & Dangoudoubiyam, S., Dec 2024, In: Parasites and Vectors. 17, 1, 426.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

Understanding temporal and spatial distribution of intestinal nematodes of horses using faecal egg counts and DNA metabarcoding

Abbas, G., Ghafar, A., Beasley, A., Stevenson, M. A., Bauquier, J., Koehler, A. V., Wilkes, E. J. A., McConnell, E., El-Hage, C., Carrigan, P., Cudmore, L., Hurley, J., Gauci, C. G., Beveridge, I., Jacobson, C., Nielsen, M. K., Hughes, K. J. & Jabbar, A., Jan 2024, In: Veterinary Parasitology. 325, 110094.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

9 Citations (Scopus)

US-wide equine strongylid egg count data demonstrate seasonal and regional trends

Nielsen, M. K., Slusarewicz, P., Kuzmina, T. A. & Denwood, M. J., 1 May 2024, In: Parasitology. 151, 6, p. 579-586 8 p.

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Open Access

2 Citations (Scopus)

Validation of a serum ELISA test for cyathostomin infection in equines

Lightbody, K. L., Austin, A., Lambert, P. A., von Samson-Himmelstjerna, G., Jürgenschellert, L., Krücken, J., Nielsen, M. K., Sallé, G., Reigner, F., Donnelly, C. G., Finno, C. J., Walshe, N., Mulcahy, G., Housby-Skeggs, N., Grice, S., Geyer, K. K., Austin, C. J. & Matthews, J. B., Jan 2024, In: International Journal for Parasitology. 54, 1, p. 23-32 10 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

10 Citations (Scopus)

Worm control practices used by Thoroughbred horse managers in Australia: A national survey

Abbas, G., Bauquier, J., Beasley, A., Jacobson, C., El-Hage, C., Wilkes, E. J. A., Carrigan, P., Cudmore, L., Hurley, J., Beveridge, I., Nielsen, M. K., Hughes, K. J., Stevenson, M. A. & Jabbar, A., Apr 2024, In: Veterinary Parasitology. 327, 110116.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

1 Citation (Scopus)

2023

Apparent treatment failure of praziquantel and pyrantel pamoate against anoplocephalid tapeworms

Nielsen, M. K., Aug 2023, In: International Journal for Parasitology: Drugs and Drug Resistance. 22, p. 96-101 6 p.

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Open Access

12 Citations (Scopus)

Assessment of worm control practices recommended by equine veterinarians in Australia

Abbas, G., Stevenson, M. A., Bauquier, J., Beasley, A., Jacobson, C., El-Hage, C., Wilkes, E. J. A., Carrigan, P., Cudmore, L., Hurley, J., Beveridge, I., Nielsen, M. K., Hughes, K. J. & Jabbar, A., 2023, In: Frontiers in Veterinary Science. 10, 1305360.

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Open Access

4 Citations (Scopus)

A statistical framework for calculating prospective sample sizes and classifying efficacy results for faecal egg count reduction tests in ruminants, horses and swine

Denwood, M. J., Kaplan, R. M., McKendrick, I. J., Thamsborg, S. M., Nielsen, M. K. & Levecke, B., Feb 2023, In: Veterinary Parasitology. 314, 109867.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

34 Citations (Scopus)

Biological implications of long-term anthelmintic treatment: what else besides resistance are we selecting for?

Nielsen, M. K., Kaplan, R. M., Abbas, G. & Jabbar, A., Nov 2023, In: Trends in Parasitology. 39, 11, p. 945-953 9 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

7 Citations (Scopus)

Egg reappearance periods of anthelmintics against equine cyathostomins: The state of play revisited

Macdonald, S. L., Abbas, G., Ghafar, A., Gauci, C. G., Bauquier, J., El-Hage, C., Tennent-Brown, B., Wilkes, E. J. A., Beasley, A., Jacobson, C., Cudmore, L., Carrigan, P., Hurley, J., Beveridge, I., Hughes, K. J., Nielsen, M. K. & Jabbar, A., Apr 2023, In: International Journal for Parasitology: Drugs and Drug Resistance. 21, p. 28-39 12 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

Open Access

17 Citations (Scopus)

Molecular diagnostics for gastrointestinal helminths in equids: Past, present and future

Ghafar, A., Abbas, G., Beasley, A., Bauquier, J., Wilkes, E. J. A., Jacobson, C., McConnell, E., El-Hage, C., Carrigan, P., Cudmore, L., Tennent-Brown, B., Hurley, J., Nielsen, M. K., Gauci, C. G., Beveridge, I., Hughes, K. J. & Jabbar, A., Jan 2023, In: Veterinary Parasitology. 313, 109851.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

10 Citations (Scopus)

Occurrence and control of equine strongyle nematode infections in Prince Edward Island, Canada

Stoughton, W. B., Begin, S., Outman, S., Stryhn, H., Yu, J., Conboy, G. & Nielsen, M. K., May 2023, In: Veterinary Parasitology: Regional Studies and Reports. 40, 100856.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

2 Citations (Scopus)

Parascaris spp. eggs shedding patterns in juvenile horses

Ripley, N. E., Gravatte, H. S., Britton, L. N., Davis, S. M., Perrin, G. M., Warner, S., Rexroat, E. K., Vetter, A. L., Maron, E. E. S., Finnerty, C. A., Stanton, V. & Nielsen, M. K., Oct 2023, In: Veterinary Parasitology. 322, 110029.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Prevalence and diversity of ascarid and strongylid nematodes in Australian Thoroughbred horses using next-generation sequencing and bioinformatic tools

Abbas, G., Ghafar, A., Bauquier, J., Beasley, A., Ling, E., Gauci, C. G., El-Hage, C., Wilkes, E. J. A., McConnell, E., Carrigan, P., Cudmore, L., Hurley, J., Beveridge, I., Nielsen, M. K., Stevenson, M. A., Jacobson, C., Hughes, K. J. & Jabbar, A., Nov 2023, In: Veterinary Parasitology. 323, 110048.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

14 Citations (Scopus)

Shortened strongylid egg reappearance periods in horses following macrocyclic lactone administration – The impact on parasite dynamics

Nielsen, M. K., Leathwick, D. M. & Sauermann, C. W., Aug 2023, In: Veterinary Parasitology. 320, 109977.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

4 Citations (Scopus)

World Association for the Advancement of Veterinary Parasitology (W.A.A.V.P.) guideline for diagnosing anthelmintic resistance using the faecal egg count reduction test in ruminants, horses and swine

Kaplan, R. M., Denwood, M. J., Nielsen, M. K., Thamsborg, S. M., Torgerson, P. R., Gilleard, J. S., Dobson, R. J., Vercruysse, J. & Levecke, B., Jun 2023, In: Veterinary Parasitology. 318, 109936.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

Open Access

85 Citations (Scopus)

2022

Anthelmintic resistance in equine nematodes: Current status and emerging trends

Nielsen, M. K., Dec 2022, In: International Journal for Parasitology: Drugs and Drug Resistance. 20, p. 76-88 13 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

Open Access

65 Citations (Scopus)

Comparison of the host response to larvicidal and nonlarvicidal treatment of naturally acquired cyathostomin infections in horses

Steuer, A. E., Scoggin, K., Stewart, J. C., Barker, V. D., Adams, A. A., Loynachan, A. T. & Nielsen, M. K., Oct 2022, In: Parasite Immunology. 44, 10, e12941.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

1 Citation (Scopus)

Efficient and Scalable Process to Produce Novel and Highly Bioactive Purified Cytosolic Crystals from Bacillus thuringiensis

Chicca, J., Cazeault, N. R., Rus, F., Abraham, A., Garceau, C., Li, H., Atwa, S. M., Flanagan, K., Soto, E. R., Morrison, M. S., Gazzola, D., Hu, Y., Liu, D. R., Nielsen, M. K., Urban, J. F., Ostroff, G. R. & Aroian, R. V., Aug 2022, In: Microbiology Spectrum. 10, 4

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

5 Citations (Scopus)

Equine strongylids: Ivermectin efficacy and fecal egg shedding patterns

Nielsen, M. K., Littman, B. A., Orzech, S. W. & Ripley, N. E., Jun 2022, In: Parasitology Research. 121, 6, p. 1691-1697 7 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

14 Citations (Scopus)

Evaluation of risk factors affecting strongylid egg shedding on Hungarian horse farms

Joó, K., Trúzsi, R. L., Kálmán, C. Z., Ács, V., Jakab, S., Bába, A. & Nielsen, M. K., Jan 2022, In: Veterinary Parasitology: Regional Studies and Reports. 27, 100663.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

8 Citations (Scopus)

How to publish a great scientific paper – A guide for publishing successfully in Veterinary Parasitology

Reichel, M. P., Greer, A. W., Nielsen, M. K. & de Waal, T., Apr 2022, In: Veterinary Parasitology. 304, 109697.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Editorial › peer-review

1 Citation (Scopus)

Label-free quantitative proteomics and immunoblotting identifies immunoreactive and other excretory-secretory (E/S) proteins of Anoplocephala perfoliata

Hautala, K., Pursiainen, J., Näreaho, A., Nyman, T., Varmanen, P., Sukura, A., Nielsen, M. K. & Savijoki, K., 16 Nov 2022, In: Frontiers in Immunology. 13, 1045468.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

3 Citations (Scopus)

Occurrence of Strongylid Nematode Parasites on Horse Farms in Berlin and Brandenburg, Germany, With High Seroprevalence of Strongylus vulgaris Infection

Jürgenschellert, L., Krücken, J., Bousquet, E., Bartz, J., Heyer, N., Nielsen, M. K. & von Samson-Himmelstjerna, G., 10 Jun 2022, In: Frontiers in Veterinary Science. 9, 892920.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

15 Citations (Scopus)

Parasite dynamics in untreated horses through one calendar year

Steuer, A. E., Anderson, H. P., Shepherd, T., Clark, M., Scare, J. A., Gravatte, H. S. & Nielsen, M. K., Dec 2022, In: Parasites and Vectors. 15, 1, 50.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

18 Citations (Scopus)

Parasite faecal egg counts in equine veterinary practice

Nielsen, M. K., Nov 2022, In: Equine Veterinary Education. 34, 11, p. 584-591 8 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

14 Citations (Scopus)

Precision of cyathostomin luminal worm counts: Investigation of storage duration and fixative

Smith, M. A., Carpenter, A. B. & Nielsen, M. K., Sept 2022, In: Veterinary Parasitology. 309, 109773.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Reflecting on the 11th International Equine Infectious Diseases Conference

Kydd, J., Nielsen, M. & Waller, A., 1 Sept 2022, In: Veterinary Record. 191, 5, p. no

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Reflections and future directions for continued development and refinement of guidelines for anthelmintic efficacy studies

Geurden, T., Smith, E. R., Vercruysse, J., Yazwinski, T., Rehbein, S. & Nielsen, M. K., Jul 2022, In: Veterinary Parasitology. 307-308, 109741.

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Open Access

2 Citations (Scopus)

Shortened egg reappearance periods of equine cyathostomins following ivermectin or moxidectin treatment: morphological and molecular investigation of efficacy and species composition

Nielsen, M. K., Steuer, A. E., Anderson, H. P., Gavriliuc, S., Carpenter, A. B., Redman, E. M., Gilleard, J. S., Reinemeyer, C. R. & Poissant, J., Nov 2022, In: International Journal for Parasitology. 52, 12, p. 787-798 12 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

35 Citations (Scopus)

The equine ascarids: resuscitating historic model organisms for modern purposes

Cain, J. L. & Nielsen, M. K., Oct 2022, In: Parasitology Research. 121, 10, p. 2775-2791 17 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Review › Research › peer-review

Open Access

6 Citations (Scopus)

The microbial community associated with Parascaris spp. infecting juvenile horses

Cain, J. L., Norris, J. K., Ripley, N. E., Suri, P., Finnerty, C. A., Gravatte, H. S. & Nielsen, M. K., Dec 2022, In: Parasites and Vectors. 15, 1, 408.

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Open Access

2 Citations (Scopus)

World association for the advancement of veterinary parasitology (WAAVP): Third edition of guideline for evaluating the efficacy of equine anthelmintics

Nielsen, M. K., von Samson-Himmelstjerna, G., Kuzmina, T. A., van Doorn, D. C. K., Meana, A., Rehbein, S., Elliott, T. & Reinemeyer, C. R., Mar 2022, In: Veterinary Parasitology. 303, 109676.

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Open Access

26 Citations (Scopus)

World association for the advancement of veterinary parasitology (WAAVP) guideline for the evaluation of the efficacy of anthelmintics in food-producing and companion animals: general guidelines

Geurden, T., Smith, E. R., Vercruysse, J., Yazwinski, T., Settje, T. & Nielsen, M. K., Apr 2022, In: Veterinary Parasitology. 304, 109698.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

24 Citations (Scopus)

2021

An inactivated bacterium (paraprobiotic) expressing Bacillus thuringiensis Cry5B as a therapeutic for Ascaris and Parascaris spp. infections in large animals

Urban, J. F., Nielsen, M. K., Gazzola, D., Xie, Y., Beshah, E., Hu, Y., Li, H., Rus, F., Flanagan, K., Draper, A., Vakalapudi, S., Li, R. W., Ostroff, G. R. & Aroian, R. V., Jun 2021, In: One Health. 12, 100241.

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Open Access

17 Citations (Scopus)

A repeatable and quantitative DNA metabarcoding assay to characterize mixed strongyle infections in horses

Poissant, J., Gavriliuc, S., Bellaw, J., Redman, E. M., Avramenko, R. W., Robinson, D., Workentine, M. L., Shury, T. K., Jenkins, E. J., McLoughlin, P. D., Nielsen, M. K. & Gilleard, J. S., Feb 2021, In: International Journal for Parasitology. 51, 2-3, p. 183-192 10 p.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

44 Citations (Scopus)

Comparative studies on faecal egg counting techniques used for the detection of gastrointestinal parasites of equines: A systematic review

Ghafar, A., Abbas, G., King, J., Jacobson, C., Hughes, K. J., El-Hage, C., Beasley, A., Bauquier, J., Wilkes, E. J. A., Hurley, J., Cudmore, L., Carrigan, P., Tennent-Brown, B., Nielsen, M. K., Gauci, C. G., Beveridge, I. & Jabbar, A., Jan 2021, In: Current Research in Parasitology and Vector-Borne Diseases. 1, 100046.

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Open Access

21 Citations (Scopus)

Cyathostomin resistance to moxidectin and combinations of anthelmintics in Australian horses

Abbas, G., Ghafar, A., Hurley, J., Bauquier, J., Beasley, A., Wilkes, E. J. A., Jacobson, C., El-Hage, C., Cudmore, L., Carrigan, P., Tennent-Brown, B., Gauci, C. G., Nielsen, M. K., Hughes, K. J., Beveridge, I. & Jabbar, A., Dec 2021, In: Parasites and Vectors. 14, 1, 597.

Research output: Contribution to journal/Conference contribution in journal/Contribution to newspaper › Journal article › Research › peer-review

Open Access

33 Citations (Scopus)

Martin Krarup Nielsen

Research output

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